- Post-doctoral Researcher, Imperial College of Science, Technology and Medicine, University of London, U.K., 1996.
- Ph.D. in Molecular Genetics, Imperial College of Science, Technology and Medicine, University of London, U.K., 1995.
- M.Sc., Medical Biology, Hacettepe University, Faculty of Medicine Ankara, Turkey, 1988.
- B.Sc., Biology, Hacettepe University, Faculty of Science, Ankara, Turkey, 1985.
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